Reproduction and Development
Male Reproductive Anatomy
In most mammalian species
The internal organs
36.1
The male’s external reproductive organs are the scrotum and penis Consist of the gonads, which produce sperm and hormones, and accessory glands
Reproductive anatomy of the human male
Seminal vesicle (behind bladder)
(Urinary bladder)
Prostate gland Urethra Scrotum
36.1
Glans penis
Bulbourethral gland Erectile tissue of penis Vas deferens Epididymis Testis
Reproductive anatomy of the human male
(Urinary bladder)
Seminal vesicle (Rectum)
(Pubic bone) Erectile tissue of penis
Vas deferens Ejaculatory duct Prostate gland Bulbourethral gland
Urethra Vas deferens Epididymis Testis
36.1
Scrotum
Glans penis Prepuce
Testes
The male gonads, or testes
The tubes are seminiferous tubules
36.1
Consist of many highly coiled tubes surrounded by several layers of connective tissue
Where sperm form
Sperm Production
Production of normal sperm
36.1
Cannot occur at the body temperatures of most mammals so the testis are held outside the body
Ducts
From the seminiferous tubules of a testis
The sperm pass into the coiled tubules of the epididymis
During ejaculation
36.1
Sperm are propelled through the muscular vas deferens, the ejaculatory duct, and exit the penis through the urethra
Glands
Three sets of accessory glands
Add secretions to the semen, the fluid that is ejaculated
A pair of seminal vesicles
36.1
Contributes about 60% of the total volume of semen
Glands
The prostate gland
The bulbourethral gland
36.1
Secretes a clear fluid directly into the urethra through several small ducts Secretes a clear mucus before ejaculation that neutralizes acidic urine remaining in the urethra
Semen in the Female Reproductive Tract
Once in the female reproductive tract
36.1
A number of processes, including contractions of the uterus, help move the sperm up the uterus
Penis
The human penis
Is composed of three cylinders of spongy erectile tissue
During sexual arousal
36.1
The erectile tissue fills with blood from the arteries, causing an erection
Female Reproductive Anatomy
The female external reproductive structures include
The internal organs are a pair of gonads
36.2
The clitoris Two sets of labia
And a system of ducts and chambers that carry gametes and house the embryo and fetus
Uterus Oviduct
(Urinary bladder) (Pubic bone)
Ovary
Reproductive anatomy of the human female (Rectum) Cervix Vagina
Urethra Shaft Glans
Bartholin’s gland
Figure 46.9
36.2
Clitoris
Prepuce
Vaginal opening
Labia minora Labia majora
Ovaries
Uterus
Oviduct
Uterine wall Endometrium
Cervix
Vagina
36.2
Follicles Corpus luteum
Ovaries
The female gonads, the ovaries
Each ovary
Is enclosed in a tough protective capsule and contains many follicles
A follicle
36.2
Lie in the abdominal cavity
Consists of one egg cell surrounded by one or more layers of follicle cells
Ovulation
The process of ovulation
The remaining follicular tissue then grows within the ovary
36.2
Expels an egg cell from the follicle
To form a solid mass called the corpus luteum, which secretes hormones, depending on whether or not pregnancy occurs
Oviducts and Uterus
The egg cell is released into the abdominal cavity
Cilia in the tube
36.2
Near the opening of the oviduct, or fallopian tube Convey the egg to the uterus
Vagina and Vulva
The vagina is a thin-walled chamber
The vagina opens to the outside at the vulva
36.2
That is the repository for sperm during copulation That serves as the birth canal through which a baby is born
Which includes the hymen, vestibule, labia minora, labia majora, and clitoris
The Reproductive Cycles of Females
In females
36.3
The secretion of hormones and the reproductive events they regulate are cyclic
The Reproductive Cycles of Females
36.3
The Ovarian Cycle
In the ovarian cycle
Hormones stimulate follicle growth, which results in ovulation
Following ovulation
The follicular tissue left behind transforms into the corpus luteum
36.4
Produces estrogen and progesterone Causes endometrial thickening
The Uterine (Menstrual) Cycle
Cycle after cycle
The maturation and release of egg cells from the ovary are integrated with changes in the uterus
If an embryo does not implant in the endometrium
36.4
A new menstrual flow begins
Conception, Embryonic Development, and Birth
In humans and most other placental mammals
36.5
Pregnancy, or gestation, is the condition of carrying one or more embryos in the uterus
Fertilization of an egg by a sperm, conception occurs in the oviduct 3 Cleavage (cell division) begins in the oviduct as the embryo is moved toward the uterus by peristalsis and the movements of cilia.
4 Cleavage continues. By the time the embryo reaches the uterus, it is a ball of cells. It floats in the uterus for several days, nourished by endometrial secretions. It becomes a blastocyst.
Ovary
2 Fertilization occurs. A sperm enters the oocyte; meiosis of the oocyte finishes; and the nuclei of the ovum and sperm fuse, producing a zygote.
5 The blastocyst implants in the endometrium about 7 days after conception.
Uterus
1 Ovulation releases a secondary oocyte, which enters the oviduct. (a) From ovulation to implantation Endometrium
Endometrium
Inner cell mass
Cavity
Blastocyst
36.5
(b) Implantation of blastocyst
Trophoblast
In fertilization, the cortical reaction modifies the zona pellucida to block polyspermy 1 The sperm migrates through the coat of follicle cells and binds to receptor molecules in the zona pellucida of the egg. (Receptor molecules are not shown here.)
2 This binding induces the acrosomal reaction, in which the sperm releases hydrolytic enzymes into the zona pellucida.
3 Breakdown of the zona pellucida 4 The nucleus and other by these enzymes allows the sperm components of the sperm to reach the plasma membrane cell enter the egg. of the egg. Membrane proteins of the sperm bind to receptors on the egg membrane, and the two membranes fuse.
Follicle cell
5 Enzymes released during the cortical reaction harden the zona pellucida, which now functions as a block to polyspermy. Zona pellucida Egg plasma membrane
Sperm basal Sperm Cortical granules body nucleus
Acrosomal vesicle EGG CYTOPLASM
After fertilization
The zygote undergoes cleavage and develops into a blastocyst before implantation in the endometrium
First Trimester
Human gestation
Can be divided into three trimesters of about three months each
The first trimester
36.6
Is the time of most radical change for both the mother and the embryo
During its first 2 to 4 weeks of development
The embryo obtains nutrients directly from the endometrium
Meanwhile, the outer layer of the blastocyst
36.6
Mingles with the endometrium and eventually forms the placenta
Blood from the embryo
Travels to the placenta through arteries of the umbilical cord and returns via the umbilical vein
Maternal arteries
Maternal veins
Placenta
Maternal portion of placenta Umbilical cord Chorionic villus containing fetal capillaries
Fetal portion of placenta (chorion)
Maternal blood pools Uterus
36.6
Fetal arteriole Fetal venule Umbilical cord
Umbilical arteries Umbilical vein
The first trimester is the main period of organogenesis
The development of the body organs
(a) 5 weeks. Limb buds, eyes, the (b) 14 weeks. Growth and heart, the liver, and rudiments development of the offspring, of all other organs have started now called a fetus, continue to develop in the embryo, which during the second trimester. is only about 1 cm long. This fetus is about 6 cm long.
36.6
(c) 20 weeks. By the end of the second trimester (at 24 weeks), the fetus grows to about 30 cm in length.
Second Trimester
During the second trimester
36.6
The fetus grows and is very active The mother may feel fetal movements The uterus grows enough for the pregnancy to become obvious
Third Trimester
During the third trimester
36.6
The fetus continues to grow and fills the available space within the embryonic membranes
A complex interplay of local regulators and hormones
Induces and regulates labor, the process by which childbirth occurs Estrogen from ovaries
Oxytocin from fetus and mother's posterior pituitary
Induces oxytocin receptors on uterus Stimulates uterus to contract Stimulates placenta to make Prostaglandins
36.7
Stimulate more contractions of uterus
Positive feedback
Birth, or parturition
Is brought about by a series of strong, rhythmic uterine contractions
The process of labor has three stages
Placenta Umbilical cord Uterus Cervix 1
Dilation of the cervix
2
Expulsion: delivery of the infant Uterus Placenta (detaching) Umbilical cord
36.8
3
Delivery of the placenta