Pla nt Nu tr iti on
Plants require certain chemical elements to complete their life cycle Plants derive most of their organic mass from the CO2 of air But they also depend on soil nutrients such as water and minerals CO2, the source of carbon for Photosynthesis, diffuses into leaves from the air through stomata.
H2O
CO2
O2
Through stomata, leaves expel H2O and O2.
O2
Minerals Roots absorb H2O and minerals from the soil.
CO2 H2O
Roots take in O2 and expel CO2. The plant uses O2 for cellular respiration but is a net O2 producer.
Ma cro nutrients a nd Mic ronutr ie nts
More than 50 chemical elements
Have been identified among the inorganic substances in plants, but not all of these are essential
A chemical element is considered essential If it is required for a plant to complete a life cycle
Nine of the essential elements are called macronutrients Because plants require them in relatively large amounts
The remaining eight essential elements are known as micronutrients Because plants need them in very small amounts
Co mmon Deficie ncies nitrogen, potassium, and phosphorus Healthy
Phosphate-deficient
Potassium-deficient
Nitrogen-deficient
Nit rogen F ix ation Nitrogen is often the mineral that has the greatest effect on plant growth Plants require nitrogen as a component of Proteins, nucleic acids, chlorophyll, and other important organic molecules
Soi l B acteri a and Nitrogen Avai labil ity Nitrogen-fixing bacteria convert atmospheric N2 to nitrogenous minerals that plants can absorb as a nitrogen source for organic synthesis Atmosphere N2
N2
Atmosphere Soil
N2
Nitrogen-fixing bacteria
Denitrifying bacteria
H (From soil) +
Soil
NH4+
NH3 (ammonia) NH4+ (ammonium)
Nitrate and nitrogenous organic compounds exported in xylem to shoot system
Nitrifying bacteria
NO3– (nitrate)
Ammonifying bacteria
Organic material (humus)
Root
So il Soil quality is a major determinant of plant distribution and growth Along with climate The major factors determining whether particular plants can grow well in a certain location are the texture and composition of the soil
Texture Is the soil’s general structure
Composition Refers to the soil’s organic and inorganic chemical components
Te xtu re a nd Co mposit ion of So il s Various sizes of particles derived from the breakdown of rock are found in soil Along with organic material (humus) in various stages of decomposition
The eventual result of this activity is topsoil A mixture of particles of rock and organic material
The topsoil and other distinct soil layers, or horizons Are often visible in vertical profile where there is a road cut or deep hole The A horizon is the topsoil, a mixture of broken-down rock of various textures, living organisms, and decaying organic matter. A
Figure 37.5
B
The B horizon contains much less organic matter than the A horizon and is less weathered.
C
The C horizon, composed mainly of partially broken-down rock, serves as the “parent” material for the upper layers of soil.
After a heavy rainfall, water drains away from the larger spaces of soil But smaller spaces retain water because of its attraction to surfaces of clay and other particles
The film of loosely bound water Soil particle surrounded by
Is usually available to plants film of water
Root hair Water available to plant
Air space (a) Soil water. A plant cannot extract all the water in the soil because some of it is tightly held by hydrophilic soil particles. Water bound less tightly to soil particles can be absorbed by the root.
Acids derived from roots contribute to a plant’s uptake of minerals When H+ displaces mineral cations from clay particles Soil particle K+
– –
Cu2+
–
– K+
–
–
–
– + K – Ca2+
Mg2+
H+ H2O + CO2
H2CO3
Root hair
HCO3 + H+ –
Cation exchange in soil. Hydrogen ions (H+) help make nutrients available by displacing positively charged minerals (cations such as Ca2+) that were bound tightly to the surface of negatively charged soil particles. Plants contribute H+ by secreting it from root hairs and also by cellular respiration, which releases CO2 into the soil solution, where it reacts with H2O to form carbonic acid (H2CO3). Dissociation of this acid adds H+ to the soil solution.