Phylogeny and Systematics putting things in order
Definitions • Phylogeny (group origin) – The evolutionary history of a species or group of related species
• Systematics – The use of morphological (appearance), and molecular comparisons to infer evolutionary relationships
23.1
Determining Phylogeny • Organisms that share very similar morphologies or similar DNA sequences are likely to be more closely related than organisms with vastly different structures or sequences
Phylogenetic Tree
Analogy vs. Homology • A potential misconception in constructing a phylogeny – Is morphological similarity due to convergent evolution, called analogy, rather than shared ancestry, called homology
23.2
Carolus Linnaeus Developed • Taxonomy – the organization of organisms into categories based on a set of characteristics
• Binomial nomenclature – the two-part format of the scientific name of an organism (genus and species) – The genus is capitalized and both words are italicized in print 23.3
Hierarchical Classification • Linnaeus also introduced a system – For grouping species in increasingly broad categories Panthera Species pardus Panthera
Genus
Felidae
Family
Carnivora
Order Class Phylum
23.3/23.4
Kingdom Domain
Mammalia Chordata Animalia Eukarya
How is taxonomic classification different from phylogenetic systematics? Taxonomy
Does not consider evolutionary history
Phylogenetic Systematics
Groups organisms using specific characters
Determines evolutionary history
23.5
Species
Panthera
Family
– In branching phylogenetic trees called cladograms
Panthera Mephitis Canis Canis Lutra lutra pardus mephitis familiaris lupus (European (leopard) (striped skunk) (domestic dog) (wolf) otter)
Order
• Systematists depict evolutionary relationships
Genus
Linking Taxonomy and Phylogeny
Felidae
Mephitis
Lutra
Mustelidae
Carnivora
Canis
Canidae
• Each branch point – Represents the divergence of two species
Leopard
23.5
Domestic cat
Common ancestor
• “Deeper” branch points – Represent progressively greater amounts of divergence
Wolf
23.5
Leopard
Common ancestor
Domestic cat
23.5
Am ph ib
D ro so ph ila La nc el et Fi sh
65.5 251 Millions of years ago
Proterozoic
542
Paleozoic
– the use of cladograms to compare the evolutionary history of different groups – depicts the number of evolutionary changes with lines of different lengths
Mesozoic Cenozoic
Cladistics
ian Bi rd Hu m R an at M ou se
Cladistics