Lucanus Leaflet

A color photo-catalogue of all known African species and subspecies of Stag Beetles. This book treats the whole African continent, including Madagascar and small islands: 125 species of 16 genera from tropical Africa and North Africa are listed and figured. For all species and subspecies the detailed distribution is given, and for some species photos of habitats and living specimens in the field are included. In addition, portraits and short biographical information about entomologists and explorers who worked particularly with the African lucanid fauna are also given. The species richness of the various countries and the perspectives for further research are discussed. For each African country all the known species are listed, together with the number of endemisms. Author, year, publication, synonyms, size, type locality, abundance, remarks, and distribution are given for every taxon.The label data of all the illustrated specimens are cited, as well as the acronym of the collection where the specimen is held. Many of the type specimens from the most important entomological museums (Berlin, Pretoria, Eberswalde, London, Paris, Cape Town, Munich, Dresden, Tervuren, Vienna, Florence, Genova and Basel) are illustrated. A large amount of bibliographical references are given at the end. More then 400 color photographs of all the species and subspecies from the African continent, as well as photos of living specimens and habitats - 192 pages, luxurious hardcover edition, large format 23x31, text in English

Hermann Julius Kolbe Henry Boileau ` Séguy Eugene Bernard Benesh Sebastian Endrödy-Younga Hugues Eli Bomans Marc Desfontaine Philippe Moretto Jean Michel Maes Tetsuo Mizukami Shinya Kawai

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` Séguy, fig. 4 - Bernard Benesh, fig. 5 - Sebastian Endrödy-Younga fig. 1 - Hermann Julius Kolbe, fig. 2 - Henry Boileau, fig. 3 - Eugene

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fig. 6 - Jean Michel Maes, fig. 7 - Hugues Eli Bomans, fig. 8 - Marc Desfontaine (on left) and Philippe Moretto, fig. 9 - Tetsuo Mizukami, fig. 10 - Shinya Kawai (on right) here together with Robert Lízler (on left) and Karl Werner

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Oonotus ( Macroonotus ) rex Endrödy-Younga Publication: ENDRÖDY-YOUNGA, 1993: 34 Type locality: South Africa, Transkei, Port St. Jones Size: m 19.1, f 15.7 Rarity: very rare Remarks: only the type specimens are known. The genus Oonotus has been revised by Endrödy-Younga (1993). R. Müller (pers. comm.), who was with S. Endrödy-Younga when he collected the holotype, told us that the specimen was slowly wandering on the bark of a tree in thick forest. Distribution: South Africa (Transkei)

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fig. 17 - m, 19.1 mm, South Africa, Transkei, Port St. Jones, 2.XII.1987, Endrödy-Younga leg., TMP, HOLOTYPUS fig. 18 - f, 15.7 mm, South Africa, Transkei, Port St. Jones, 2.XII.1987, Endrödy-Younga leg., TMP, ALLOTYPUS

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Figulus sublaevis (Palisot de Beauvois) Publication: PALISOT DE BEAUVOIS, 1805: 3 (Lucanus) Synonyms: anthracinus Klug, 1833: 85 ebenus Westwood, 1834: 119 nigritus Westwood, 1838: 261 monilifer Parry, 1862: 113 vulneratus J. Thomson, 1862: 433 lettowi Kriesche, 1921: 113 nossibenus Kriesche, 1921: 114 Type locality: “Regne Oware” Size: 11.4 -17.4 Rarity: common Remarks: according to Maes (1992a: 53) F. decipies is a subspecies of F. sublaevis inhabiting only Sao Thomé Island, but according to Bomans (1992: 251) and Monte & Bartolozzi (2004, in press) F. decipies is a valid species. F. anthracinus seems to be a valid species too, but further studies are required. On several occasions individuals of Figulus sublaevis were collected at light by the second author and R. Lizler in open forests of southern and eastern Africa. Distribution: Cameroon, Democratic Republic of the Congo (ex Zaire),Gabon, Guinea, Malawi, Mozambique, Madagascar, Senegal, Tanzania, Togo

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fig. 42 - 15 mm, Principe Isl., AKC fig. 43 - m, 17 mm, ex Coll. Borel, AKC fig. 44 - m, 13 mm, Tanzania, Morogoro Prov., Uluguru Mts., 8. - 9.XII.1996, Werner & Lizler leg., KWC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Figulus

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Amneidus godefroyi Coquerel Publication: COQUEREL, 1866: 326 Type locality: Is. Bourbon Size: m 13.0 -15.0, f 12.0-15.0 Rarity: rare Remarks: interesting remarks on this species are given by Paulian (1959, 1973). Distribution: La Réunion Is.

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fig. 26 - m, 15 mm, Réunion, I.1996, AKC

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fig. 27 - f, 15 mm, La Réunion, Kerveguen, 10.I.1962, AKC

Nigidius gnu Kriesche Publication: KRIESCHE, 1921: 109 Type locality: D. Ost Afrika, Pangani, Pugu c/o Daressalam Size: 13.0 -18.0 Rarity: very rare Remarks: close to N. bubalus. Bomans redescribed this species and established the lectotype. Distribution: Tanzania

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fig. 73 - 17.5 mm, D. O. Afrika, Pangani, ZMB, LECTOTYPUS

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Nigidius

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fig. 119 - Madagascar, Marojejy, March 2001 fig. 120 - Madagascar, Ankarana, March 2001 fig. 121 - Madagascar, “Andasibe (Périnet)”, May 1998

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fig. 122 - Madagascar, Morondava, April 1998 fig. 123 - Madagascar, Beza Mahafaly, April 1998 fig. 124 - Madagascar, Ranomafana, March 1998 fig. 125 - Madagascar, Ranomafana, March 1998

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Ganelius

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Dendezia renieri Basilewsky Publication: BASILEWSKY, 1952: 37 Type locality: Rwanda, Dendezi Size: m 23.0 -25.0, f [size not indicated in the description] Rarity: very rare Remarks: this species is very peculiar for the presence of a medial cephalic horn. We are grateful to A. Drumont (IRSNB) for the fine pictures of this rare species. B. Gill (pers. comm.) found a dead specimen of Dendezia renieri on a dirt road in the Kibale forest of Uganda in October, 2001. Distribution: SE Democratic Republic of the Congo (ex Zaire), Rwanda, Uganda

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fig. 126 - m, 25 mm, Ruanda, Ihembe, 11 km E de Dendezi, 2000km, 19.IX.1951, H. RENIER leg, IRSNB, PARATYPUS fig. 127 - m, 24 mm, Zaire, Kahuzi, km 82, 1-15.IX.1951, legit H. Bomans (in coll. G. Taroni)

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Colophon berrisfordi Barnard Publication: BARNARD, 1932: 173 Type locality: South Africa, Zwartberg Range, Meiringspoort Berg Size: m 19.0 - 22.0 Rarity: very rare Remarks: the genus Colophon has been revised by Endrödy-Younga (1988) and Mizukami & Kawai (1997). R. Lizler (pers. comm.) collected a single male of C. berrisfordi among several specimens of C. whitei under stones. Distribution: South Africa (Swartberg)

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fig. 137 - m, 19 mm, Rep. of S. Africa, Cape, Swartberg, IX.1993, Lizler leg., AKC

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Colophon cassoni Barnard Publication: BARNARD, 1932: 172 Type locality: South Africa, Zwartberg Range, Meiringspoort Berg Size: m 16.0 - 24.0, f 16.0 - 17.0 Rarity: very rare Remarks: the genus Colophon has been revised by Endrödy-Younga (1988) and Mizukami & Kawai (1997). This species is active only during proper weather conditions. Distribution: South Africa (Swartberg)

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fig. 140 - m, 17.5 mm, Rep. of S. Africa, Cape, Meiringspoortberg, I.1994, AKC fig. 141 - f, 16 mm, Rep. of S. Africa, Cape, Meiringspoortberg, I.1994, AKC

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fig. 142 - Swartberg Mts., Rep. of S. Africa, March 1993 fig. 143 - C. haughtoni - m, Rep. of S. Africa, Cape, Hexriver Mts., Matroosberg, I. 1993 fig. 144 - C. eastmani nagaii - m, Rep. of S. Africa, Cape Province, Langeberg, Dassie'shoek, 29. I. 1993, fig. 145 & 146 - C. izardi - m, Rep. of S. Africa, Cape Province, Langeberg, Boosmansbos, 27. XII. 1992

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Colophon

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Prosopocoilus kasaiensis Maes Publication: MAES, 1990: 13 Synonym: swanzianus sensu Auctorum (partim) Type locality: Congo Belge, Dekese, Kasai Size: m 28.0 - 32.0, f 20.0 - 22.0 Rarity: very rare Remarks: this species was described as a form of P. swanzianus, then elevated to subspecies by Maes (1992b); according to Desfontaine & Moretto (2003) it is a valid species, very close to P. demeyeri, and can be separated only by the examination of the genitalia. Distribution: Democratic Republic of the Congo (ex Zaire)

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fig. 268 - m, 32 mm, Congo Belge, Kasai, Kondue, Leohard leg., MRAC, PARATYPUS fig. 269 - m, 28 mm, Congo Belge, Kasai, Kondue, Leohard leg., MRAC, PARATYPUS

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Prosopocoilus katanganus Bomans Publication: BOMANS, 1967a: 392 Type locality: Rép. du Congo, Kafakumba Size: m 31.0 - 34.5, f 20.0 - 26.0 Rarity: very rare Remarks: this species is close to P. congoensis. Distribution: Cameroon, Democratic Republic of the Congo (ex Zaire)

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fig. 270 - m, 34.5 mm, Kamerun, 1968, AKC fig. 271 - f, 26 mm, Kamerun, 1968, AKC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Prosopocoilus

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Prosopocoilus umhangi (Fairmaire) Publication: FAIRMAIRE, 1891b: 192 (Cladognathus) Synonyms: hacquarti Oberthür, 1880: 119 wentzelheckmannae Kolbe, 1901: 71 (Metopodontus) elizabethae Waterhouse, 1905: 375 (Metopodontus) Type locality: Zanzibar Size: m 28.2 - 69.1, f 26.0 - 38.0 Rarity: not rare Remarks: P. umhangi specimens were collected in numbers by the second author and by R. Lizler on trees of rain forest in central Tanzania. Distribution: Malawi, Tanzania

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fig. 334 - m, 69 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC fig. 335 - f, 38 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC

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fig. 336 - m, 34 mm, Tanzania, Usambara, AKC fig. 337 - f, 26 mm, Tanzania, Usambara, AKC fig. 338 - m, 37 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC fig. 339 - m, 67 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC fig. 340 - f, 32.5 mm, D.O.A., Nkonda Uguro, 1910, AKC fig. 341 - m, 57 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Prosopocoilus

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Lucanus tetraodon Thunberg Publication: THUNBERG, 1806: 188 Synonyms: for the long list of synonyms see Krajcik (2001) Type locality: Italy Size: m 26.1- 50.0, f 25.8 - 37.2 Rarity: common Remarks: L. tetraodon is a palaearctic species occurring in oak forests. In North Africa the species is quoted only from Algeria. Distribution: Algeria; Balkan Peninsula, France (including Corsica), S Italy (including Sicily)

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fig. 360 - m, 45 mm, Italia, Basilicata, Gallipoli-Cognato, 7.VII.1997, B. Cecchi & B. Carletti leg., LBC fig. 361 - f, 32 mm, Sicilia, San Fratello, Messina Prov., 1.VII.1984, Aliquó leg., LBC

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fig. 362 - m, 38 mm, Italia, Calabria, Capistrano, Catanzaro Prov., 800 – 1000 m, VIII.1990, I. Bonato leg., LBC fig. 363 - m, 28 mm, Italia, Calabria, Capistrano, Catanzaro Prov., 800 – 1000 m, VIII.1990, I. Bonato leg., LBC fig. 364 - m, 42 mm, Sicilia, Messina Prov., P. Femmina Morta, 9.VII.1983, Aliquó leg., LBC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae)

Lucanus

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