Transport and Circulation
We’ll be discussing 1. Cellular Transport 2. Plant Tissues and Transport in Plants 3. Trends and Various Strategies Used by Animals to Transport Materials 4. Transport in Man 5. Disorders of the Circulatory System
Membranes and cellular transport
Active vs Passive Transport
Transport of large molecules
Summary of Transport Processes Diffusion
Direction
Osmosis
Facilitated Diffusion
Pumps
High to Low High to Low High to Low Low to High
Endocytosis /exocytosis N/A
Transport Mechanism
Pores
Pores
Channels
Pumps
Membrane
Energy Required?
No
No
No
Yes
Yes
Type of particle
Small, nonpolar
Water
Smallmedium
Ions
Small-large
Examples of particles
CO2, O2
H2O
Glucose, fructose, Na+, Ca+2
Na+, K+, H+
Food, waste
Transport within the eukaryotic cell
Endomembrane system • Endoplasmic reticulum – manufacturing and transport facility – proteins produced in rough ER are packaged in vesicles
• Golgi apparatus – modification and storage facility – receiving end and shipping end
• Vacuole – large membrane bound sacs – usually stores undigested nutrients
Cyclosis/cytoplasmic streaming
TRANSPORT IN PLANTS
VASCULAR TISSUES: XYLEM AND PHLOEM Xylem
Tracheids* Vessel elements* Parenchyma cells Fiber
Phloem
Sieve-tube members Companion cells Sclerenchyma fibers Parenchyma cells
Both
are continuous throughout the plant body
TRANSPORT OCCURS ON THREE LEVELS 1.
2.
3.
Uptake and release of water and solutes by individual cells Short-distance transport of substances by tissues and organs Long-distance transport of minerals in water and sap within xylem and phloem by the whole plant body
A VARIETY OF PHYSICAL PROCESSES ARE INVOLVED IN THE DIFFERENT TYPES OF TRANSPORT
4 Through stomata, leaves take in CO2 and expel O2. The CO2 provides carbon for photosynthesis. Some O2 produced by photosynthesis is used in cellular respiration.
CO2
O2
5 Sugars are produced by photosynthesis in the leaves.
Light H2O
Sugar
3 Transpiration, the loss of water
from leaves (mostly through stomata), creates a force within leaves that pulls xylem sap upward.
6
Sugars are transported as phloem sap to roots and other parts of the plant.
Water and minerals are transported upward from roots to shoots as xylem sap. 2
1 Roots absorb water and dissolved minerals from soil.
O2 H2O Minerals
CO2
7 Roots exchange gases with the air spaces of soil, taking in O2 and discharging CO2. In cellular respiration, O2 supports the breakdown of sugars.
TRANSPORT OF IONS AT THE CELLULAR LEVEL DEPENDS ON SELECTIVELY PERMEABLE MEMBRANES
Controls the movement of solutes into and out of the cell With specific transport proteins Enable plant cells to maintain an internal environment different from their surroundings
SHORT-DISTANCE H2O TRANSPORT IN THE ROOT
WATER AND MINERALS ASCEND FROM ROOTS TO SHOOTS THROUGH THE XYLEM ROOT PRESSURE TRANSPIRATION–COHESION– TENSION THEORY
Tension – negative pressure
STOMATA HELP
Cuticle Upper epidermal tissue
REGULATE THE RATE OF TRANSPIRATION
Leaves – broad surface areas Increase photosynthesis Increase water loss through stomata (transpiration)
Turgid
Lower epidermal Trichomes Stomata tissue (“hairs”)
Flaccid
100 µm
Flaccid
Turgid
ORGANIC NUTRIENTS
ARE
Vessel (xylem)
Sieve tube (phloem)
Source cell (leaf)
1
TRANSLOCATED THROUGH THE PHLOEM (PRESSURE – FLOW MODEL)
H2O
Sucrose
1
H2O
Translocation – transport of organic molecules in the plant Phloem sap
2
Pressure flow
Transpiration stream
Mostly sucrose Sugar source sugar sink Source is a producer of sugar Sink is a consumer/storage facility for sugar
2
4
3 H2O
3
4 Sink cell (storage root)
Sucrose
Loading of sugar (green dots) into the sieve tube at the source reduces water potential inside the sieve-tube members. This causes the tube to take up water by osmosis.
This uptake of water generates a positive pressure that forces the sap to flow along the tube. The pressure is relieved by the unloading of sugar and the consequent loss of water from the tube at the sink. In the case of leaf-to-root translocation, xylem recycles water from sink to source.
PRESSURE – FLOW MODEL 1.
2.
3.
high solute concentration at source increase in hydrostatic pressure sugars in sink draw water out of phloem